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Jeff Otto of IGN called it "a tension-filled drama with great performances".Later that year, she landed the part of Rose, the devious—but lovable—neighbor of Charlie Harper (Charlie Sheen) on the CBS sitcom Two and a Half Men.(all 2009), Win Win (2011), and The Perks of Being a Wallflower (2012).
The romantic drama Hello I Must Be Going (2012), in which Lynskey portrayed Amy, a dispirited 30-something who finds herself having to move back in with her parents, was particularly well received by critics.
Kenneth Turan of The Los Angeles Times wrote: "If you know the name Melanie Lynskey, you're already planning to see her in Hello I Must Be Going. Club described Beatrice as "the series' standout character", noting that "Lynskey steals the show with her amazing putdowns and passive-aggressiveness, smartly avoiding overdone sass or sarcasm".
I had this double life: I had this whole indie side of my career, and people in that world didn't know I was on this huge sitcom.
Then, people would recognize me from Two and a Half Men and think that I never had another job. In 2006, she had a supporting role in Clint Eastwood's epic war drama Flags of Our Fathers, based on the book of the same name about the 1945 Battle of Iwo Jima.
The series was a ratings hit with an average of 18.5 million viewers over three consecutive nights.
In 2003, Lynskey played a writer for The New Republic in Shattered Glass, a film based on the exploits of former journalist Stephen Glass (Hayden Christensen).The latter film was nominated for six Academy Awards, including Best Picture.The following year, Lynskey appeared in Seeking a Friend for the End of the World with Steve Carell, and had a key role in the film adaptation of Stephen Chbosky's novel The Perks of Being a Wallflower, playing the mentally disturbed Aunt Helen.Wesley Morris of The Boston Globe called it "the best performance in the movie".Soderbergh told The Los Angeles Times: "She is so watchable.You never quite know what you're going to get, you just know it's going to be good.Her rhythms are really unusual, like her cadence and her reaction times to things, and the way she sort of lays out a sentence. Also in 2009, Lynskey appeared as Edward Norton's pregnant fiancée in Leaves of Grass, and in Jason Reitman's Up in the Air, where she played the sister of George Clooney.On television, Lynskey's best known role is that of Rose on the CBS sitcom Two and a Half Men (2003–15).She also appeared as Michelle Pierson on the HBO series Togetherness (2015–16), for which she received a nomination for the Critics' Choice Television Award for Best Supporting Actress in a Comedy Series.Asked by an interviewer in 2012 on how she felt about typically being cast—up to that point—as a supporting player rather than a lead, Lynskey said it was something she had thought about a lot, and that the "meaty" parts are written for men, or Meryl Streep.Speaking in 2017 about taking risks in her film work, she said: "I want to tell stories about women who are interesting and complicated and not like people you've seen before [...] There aren't that many opportunities [to do that] except in the independent film world.